Here are some quotes from the pop article:
A rocky, ocean-bearing planet with more than 0.1 percent methane in its atmosphere should be considered a potentially inhabited planet, the researchers say. And if the atmospheric methane reaches levels of 1 percent or more? In that case, “potentially” doesn’t cut it—such a world would “likely” be home to alien life.
Thus, methane and carbon dioxide together, unaccompanied by carbon monoxide, on a rocky, ocean-bearing world would best be interpreted as an airtight sign of anoxic life.
Now, some information about Titan from the Wikipedia page about Titan:
Titan's atmosphere Stratosphere: 98.4% nitrogen 1.4% methane 0.2% hydrogen Lower troposphere: 95.0% nitrogen 4.9% methane -OR, depending on who you ask- 97% nitrogen 2.7% methane
The density of Titan is consistent with a body that is about 60% rock and 40% water.
There are trace amounts of other hydrocarbons, such as [...] carbon monoxide [...].
If I understand correctly, 'trace amounts' of CO is in line with the requirements stated in the Catling paper.
So, is Titan covered in microbes, or what? Since the article didn't mention Titan, I presume I missed something. What was it?